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«CURRICULUM VITAE BENJAMIN S. TWINING Senior Research Scientist Director of Research and Education Bigelow Laboratory for Ocean Sciences 60 Bigelow ...»

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Cochlan, F.J. Millero, P.G. Falkowski, J.E. Bauer, R.H. Wanninkhof, R.M. Kudela, M.A.

Altabet, B.E. Hales, T. Takahashi, M.R. Landry, R.R. Bidigare, X. Wang, Z. Chase, P.G.

Strutton, G.E. Friederich, M.Y. Gorbunov, V.P. Lance, A.K. Hilting, M.R. Hiscock, M.

Demerest, W.T. Hiscock, K.F. Sullivan, S.J. Tanner, R.M. Gordon, C.N. Hunter, V.A.

Elrod, S.E. Fitzwater, J.L. Jones, S. Tozzi, M. Koblizek, A.E. Roberts, J. Herndon, J.

Brewster, N. Ladizinsky, G. Smith, D. Cooper, D. Timothy, S.L. Brown, K.E. Selph, C.C. Sheridan, B.S. Twining, and Z.I. Johnson. 2004. Southern Ocean Iron Enrichment Experiment (SOFeX): Carbon cycling in high- and low-Si waters. Science. 304: 408-414.

Twiss, M.R., B.S. Twining, and N.S. Fisher. 2004. Bioconcentration of inorganic and organic

thallium by freshwater phytoplankton. Environmental Toxicology and Chemistry. 23:

968-973.

Twining, B.S. and N.S. Fisher. 2004. Trophic transfer of trace metals from protozoa to mesozooplankton. Limnology & Oceanography. 49: 28-39.

Twining, B.S., S.B. Baines, N.S. Fisher, J. Maser, S. Vogt, C. Jacobsen, A. Tovar-Sanchez, S.

Sañudo-Wilhelmy. 2003. Quantifying trace elements in individual aquatic protist cells with a synchrotron x-ray fluorescence microprobe. Analytical Chemistry. 75: 3806-3816.

Twining, B.S., M.R. Twiss, and N.S. Fisher. 2003. Oxidation of thallium by Great Lakes plankton communities. Environmental Science and Technology. 37: 2720-2726.

Wong, W.H., J.S. Levinton, B.S. Twining, N.S. Fisher, B.P. Kelaher, and A.K. Alt. 2003.

Assimilation of carbon from a rotifer by mussels Mytilus edulis and Perna viridis: a potential marine food web link. Marine Ecology Progress Series. 253: 175-182.

Mylon, S.E., B.S. Twining, N.S. Fisher, and G. Benoit. 2003. Relating the speciation of Cd, Cu, and Pb in two Connecticut rivers with their uptake in algae. Environmental Science and Technology. 37: 1261-1267.

Wong, W.H., J.S. Levinton, B.S. Twining, and N.S. Fisher. 2003. Assimilation of micro- and mesozooplankton by zebra mussels: A demonstration of the food web link between zooplankton and benthic suspension feeders. Limnology & Oceanography. 48: 308-312.

Twiss, M.R., B.S. Twining, and N.S. Fisher. 2003. Partitioning of dissolved thallium by seston in Lakes Erie and Ontario. Canadian Journal of Fisheries and Aquatic Sciences. 60: 1369Twining, B.S., J.J. Gilbert, and N.S. Fisher. 2000. Evidence of homing behavior in the coral reef mysid Mysidium gracile. Limnology & Oceanography. 45: 1845-1849.

Beaulieu, S.E., M.M. Mullin, V.T. Tang, S.M. Pyne, A.L. King, and B.S. Twining. 1999. Using an optical plankton counter to determine the size distributions of preserved zooplankton samples. Journal of Plankton Research. 21: 1939-1956.

Submitted Boyd, P.W., R. Strzepek, M. Ellwood, D.A. Hutchins, S. Nodder, B.S. Twining, and S.W.

Wilhelm. 2014. Why are biotic iron pools uniform across high- and low-iron pelagic ecosystems? Global Biogeochemical Cycles. In review.

The GEOTRACES Group (135 co-authors). 2014. The GEOTRACES Intermediate Date Product

2014. Marine Chemistry. In review.

Pižeta, I., S.G. Sander, R.J.M. Hudson, O. Baars, K.A. Barbeau, K.N. Buck, R.M. Bundy, G.

Carrasco, P. L. Croot, C. Garnier, L.J.A. Gerringa, M. Gledhill, K. Hirose, Y. Kondo, L.M. Laglera, J. Nuester, D. Omanović, M.J.A. Rijkenberg, S. Takeda, B.S. Twining, M.

Wells. Quantitative analysis of complexometric titration data: An intercomparison of methods for estimating models of metal complexation by mixtures of natural ligands.

Marine Chemistry. In review.

Reports and conference proceedings De Jonge, M.D., B. Hornberger, C. Holzner, B. Twining, D. Paterson, I. McNulty, C. Jacobsen, and S. Vogt. 2008. Quantitative scanning differential phase contrast microscopy. Journal de Physique IV.

Twining, B.S., S.B. Baines, and N.S. Fisher. 2004. Metal cycling through plankton communities:

a single-cell approach using synchrotron-based x-ray fluorescence. Rapp. Comm. Int.

Mer. Medit. 37: 251.

Twining, B.S., S.B. Baines, and N.S. Fisher. 2003. Quantification of Si, P, S, Mn, Fe, and Zn in cultured phytoplankton and Southern Ocean protists. 2002 APS User Activity Report. 2 pp.

Twining, B.S., S.B. Baines, N.S. Fisher, C. Jacobsen, and J. Maser. 2003. Quantification and localization of trace metals in natural plankton cells using a synchrotron x-ray fluorescence microprobe. Journal de Physique IV. 104: 435-438.

Twining, B.S., S.B. Baines, N.S. Fisher, C. Jacobsen, and J. Maser. 2001. Quantification of trace elements in cultured phytoplankton cells using an x-ray fluorescence microprobe. 2001 APS User Activity Report. 2 pp.

Twining, B.S., S.B. Baines, and N.S. Fisher. 2001. Measurement of metal concentrations in marine nanoplankton cells using a x-ray fluorescence microprobe. Rapp. Comm. Int.

Mer. Medit. 36: 169.

Twining, B.S., S.B. Baines, N.S. Fisher, C. Jacobsen, and J. Maser. 2000. Quantification and localization of elements in nanoplankton cells using an x-ray fluorescence microprobe.

2000 APS User Activity Report. 2 pp.

Invited presentationsU

October, 2014. Decoding the ocean’s black box: exploration in global biogeochemistry through microbial oceanography. Bates College.





April, 2014. Tracking the metallomic responses of phytoplankton to environmental gradients in the ocean. EAWAG. Dubendorf, Switzerland.

October, 2013. Unraveling the roles of plankton in ocean biogeochemistry, one cell at a time.

University of Tennessee—Knoxville. Knoxville, TN.

August, 2013. The microbial ocean: Epicenter of life on earth. Phippsburg Land Trust Annual Meeting. Sebasco Harbor Resort, ME.

August, 2013. Application of synchrotron x-ray fluorescence mapping and absorption spectroscopy in marine (and environmental) science. NSLS-II Early Science Workshop.

Brookhaven National Laboratory. Upton, NY August, 2013. Studying the ocean through SXRF analyses of its smallest residents. X-ray Fluorescence Microscopy in Biology Workshop. Northwestern University. Chicago, IL.

May, 2013. Tracking the metallomic responses of phytoplankton to environmental gradients in the ocean. 96th Canadian Chemistry Conference. Québec, Quebec.

February, 2013. The microbial ocean: Epicenter of life on earth. First Parish Church.

Brunswick, ME.

December, 2012. Trace metal composition of marine plankton: Linking ecosystem form to biogeochemical function. Stony Brook University. Stony Brook, NY.

December, 2012. Silent Spring Revisited: 50 Years Later. Cornerstones of Science. Portland, ME.

November, 2011. Role of plankton in the cycling of trace elements and isotopes. GEOTRACES Data-Model Synergy Workshop. Barcelona, Spain.

October, 2011. Exploring the Southern Ocean with Maine Scientists: Bringing Oceanography to Island Classrooms. 2011 Island Institute Teacher’s Conference. Rockland, ME.

August, 2011. Trace metal composition of marine plankton: Linking ecosystem form to biogeochemical function. Chemical Oceanography Gordon Research Conference.

Andover, NH.

June, 2011. FeCycle II: Quantifying the cycling of Fe and C during a spring diatom bloom.

Modeling and Synthesis of Southern Ocean Natural Iron Fertilization Workshop. Woods Hole Oceanographic Institution. Woods Hole, MA.

April, 2011. Insights into oceanic metal cycling from single-cell element analysis. Massachusetts Institute of Technology. Cambridge, MA.

October, 2010. Studying ocean biogeochemistry, one cell at a time. Lafayette College. Easton, PA.

October, 2010. Antarctic Icebergs: floating estuaries in a warming world. COSEE Ocean Systems ROLE Model Webinar.

September, 2010. Global warming and Antarctic icebergs: floating estuaries in the Southern Ocean. Carleton-Willard Village. Bedford, MA.

June, 2010. Trace nutrient limitation and individual cell elemental composition. Bowdoin College. Brunswick, ME.

June, 2010. Mapping trace metals in marine plankton: clues to metal function and fate.

Environmental Inorganic Chemistry Gordon Research Conference. Newport, RI.

March, 2010. Studying ocean biogeochemistry, one cell at a time. Colby College. Waterville, ME.

November, 2009. Elemental composition of plankton in the equatorial Pacific Ocean: Evidence of diatom co-limitation by iron and silicic acid. Old Dominion University. Norfolk, VA.

July, 2009. Global warming and the proliferation of icebergs: Floating estuaries in the Southern Ocean. Thornton Oaks Retirement Community. Brunswick, ME.

June, 2009. Global warming and the proliferation of icebergs: Floating estuaries in the Southern Ocean. Café Scientifique. Boothbay Harbor, ME.

May, 2009. Ocean science in the 21st century: Where has ocean science come since the publication of The Sea Around Us? Cornerstones of Science. Brunswick, ME.

May, 2009. Global climate change, the role of the oceans, and icebergs in the Weddell Sea. First Church in Wenham. Wenham, MA.

February, 2009. Examining the role of plankton in ocean biogeochemistry, one cell at a time.

University of Maine. Orono, ME.

November, 2007. Examining the role of plankton in ocean biogeochemistry, one cell at a time.

Georgia Institute of Technology. Atlanta, GA.

August, 2007. Examining the elemental composition of ocean plankton one cell at a time.

University of Rhode Island Graduate School of Oceanography. Narragansett, RI.

August, 2007. The roles of protists in ocean iron biogeochemistry. Joint Annual Meeting of the International Society of Protistologists and the Phycological Society of America.

Providence, RI.

June, 2007. Unraveling the roles of plankton in ocean biogeochemistry. Bigelow Laboratory for Ocean Sciences. Boothbay Harbor, ME.

March, 2007. Phytoplankton trace metal quotas across nutrient gradients in HNLC waters. Duke University Marine Lab. Beaufort, NC.

December, 2006. Examining the elemental composition of ocean plankton one cell at a time.

University of South Carolina Department of Geology. Columbia, SC.

November, 2006. Planktonic metal stoichiometries in HNLC waters: A single-cell approach using synchrotron-based x-ray fluorescence. Texas A&M University—Galveston, TX.

October, 2006. Dynamic metal quotas in ocean phytoplankton: Insights from single-cell x-ray analysis. University of South Carolina Plant Biology Seminar. Columbia, SC.

October, 2006. Trace metal stoichiometries in ocean plankton: Insights from single-cell x-ray analysis. Coastal Georgia Environmental Symposium. Savannah, GA.

June, 2006. Metal cycling through plankton communities: a single-cell approach utilizing T synchrotron-based x-ray fluorescence. Bigelow Laboratory for Ocean Sciences.

Boothbay Harbor, ME.T May, 2006. Breakthroughs in nano-related environmental- and geo-science: Scaling from nano to global. Molecular- and nano-environmental geochemistry working group. National Science Foundation.

July, 2005. Measuring element stoichiometries of ocean plankton with a hard x-ray fluorescence microprobe. Advanced Photon Source User Science lunch seminar.

May, 2005. Measuring element stoichiometries of ocean plankton with a hard x-ray fluorescence microprobe. Advanced Photon Source Users Meeting.

February, 2005. Stoichiometric responses of Southern Ocean plankton to iron fertilization:

Expanding stoichiometry to include trace elements. American Society of Limnology and Oceanography Aquatic Sciences Meeting. Salt Lake City, UT.

August, 2004. Metal cycling through plankton communities: a single-cell approach utilizing synchrotron-based x-ray fluorescence. University of South Carolina, Department of Chemistry and Biochemistry.

April, 2004. Metal cycling through plankton communities: a single-cell approach. Yale Institute for Biospheric Studies seminar series.

May, 2003. Quantification and localization of trace metals in marine and freshwater protists using a synchrotron x-ray fluorescence microprobe. National Synchrotron Light Source Users meeting.

March, 2003. Element stoichiometries within Southern Ocean nanoplankton communities as measured with a synchrotron x-ray fluorescence microprobe. Woods Hole Oceanographic Institution, Marine Chemistry & Geochemistry.

May, 2001. Measurement of trace element concentrations in marine and freshwater microbes using x-ray microscopy. Biological applications of x-ray microbeams workshop, Advanced Photon Source.

Contributed presentations (*denotes student or post-doc presentation)U

*Ohnemus, D.O., S. Rauschenberg, and B.S. Twining. 2014. Trace element composition of phytoplankton along the US GEOTRACES Pacific Zonal Transect: Comparing singlecell SXRF quotas, chemical leaching, and bulk particle digestion. AGU Fall Meeting.

San Francisco, CA.

Twining, B.S., S. Rauschenberg, J. Vedamati, J. Moffett, and P. Sedwick. 2014. Response of phytoplankton metal quotas to oceanic gradients in dissolved and particulate metals.

Goldschmidt Conference. Sacramento, CA.

Glass, J.B., S.E. McGlynn,G. Chadwick, K.S. Dawson, S. Chen, S. Vogt, B. Lai, J. Deng, E.D.

Ingall, B.S. Twining, and V.J. Orphan. 2014. Nano-scale elemental imaging of microbes and minerals from deep sea methane seeps. Goldschmidt Conference. Sacramento, CA.

Twining, B.S., S.E.K. Rauschenberg, P.N. Sedwick, J.N. Fitzsimmons, and K.N. Buck. 2014.

Iron quotas of North Atlantic phytoplankton reflect biogeochemical environment. Ocean Sciences Meeting. Honolulu, HI.

Fields, D.M., H.I. Browman, and B.S. Twining. 2014. Copepods intestines: 1021 microbioreactors of global ocean processes. Ocean Sciences Meeting. Honolulu, HI.

Balch, W.M., N.R. Bates, B.S. Twining, P.J. Lam, and D.T. Drapeau. 2014. The Great Calcite Belt and the saga of residual nitrate. Ocean Sciences Meeting. Honolulu, HI.

*Chapman, A.U., J. Nuester, and B.S. Twining. 2014. Regeneration of Fe(II) by protist grazing in the ocean. Ocean Sciences Meeting. Honolulu, HI.

Baines, S.B., M.R. Landry, S.L. Smith, B.S. Twining, and X. Chen. 2014. Trace metal limitation of zooplankton in the Costa Rican upwelling dome. Ocean Sciences Meeting. Honolulu, HI.

*Nuester, J., S. Rauschenberg, A. Chapman, D.M. Fields, and B.S. Twining. 2014. Zooplankton grazing produces highly bioavailable iron. Ocean Sciences Meeting. Honolulu, HI.



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