WWW.DISSERTATION.XLIBX.INFO
FREE ELECTRONIC LIBRARY - Dissertations, online materials
 
<< HOME
CONTACTS



Pages:     | 1 |   ...   | 18 | 19 || 21 | 22 |   ...   | 34 |

«Taxonomic Revision, Molecular Phylogeny and Zoogeography of the huntsman spider genus Eusparassus (Araneae: Sparassidae) Dissertation for attaining ...»

-- [ Page 20 ] --

Etymology. The species is named in honour of Dr Rudy Jocqué (MRAC). The visit of the author to MRAC coincided with his retirement. The author would like to dedicate the name of this conspicuous species to him in recognition of his long time of productive arachnological research especially on African spiders; noun in genitive case.

Diagnosis. This is the only Eusparassus species whose male’s dRTA is bifurcated at its distal end (Figs 35a, b) and female epigyne with MS clearly visible posteriorly (Fig. 36a) [see also diagnosis for jaegeri species group above].

Description. Male (ranges: n=6, single measurement: holotype):

Measurements. Males medium-sized. Total length 12.0–13.8, prosoma length 5.7–6.3, prosoma width 4.6–5.4, anterior width of prosoma 2.7–3.0, opisthosoma length 6.3–7.5, opisthosoma width 4.0. Eye diameters: AME 0.47, ALE 0.33, PME 0.31, PLE 0.34; eye interdistances: AMEAME 0.18, AME-ALE 0.08, PME-PME 0.42, PME-PLE 0.45, AME-PME 0.33, ALE-PLE 0.22, clypeus height at AME 0.23, clypeus height at ALE 0.37. AME largest, other eyes subequal (Fig.

35d).

Results: chapter 3.2: Systematics and zoogeography with revision of Afro-Arabian species FIGURE 35. Eusparassus jocquei spec. nov., holotype male from Zimbabwe: Bulawayo (MRAC) (a) left palp, ventral; (b) left palp, retrolateral; (c) embolus tip and conductor, ventral; (d) eye arrangement, dorsal;

(e) right chelicera, ventral.

Results: chapter 3.2: Systematics and zoogeography with revision of Afro-Arabian species FIGURE 36. Eusparassus jocquei spec. nov., paratype female from Zimbabwe: Bulawayo (MRAC). (a) epigyne, ventral; (b) vulva, dorsal; (c) left vulva, anterio-dorso-lateral.

Chelicerae. Chelicerae with 2 anterior and 3 or 4 posterior teeth, cheliceral furrow with a line of 7 to 10 intermarginal denticles (Fig. 35e).

Results: chapter 3.2: Systematics and zoogeography with revision of Afro-Arabian species Legs. Leg formula: II I IV III. Measurements of palp and legs: Palp 8.3 [2.8, 1.2, 1.3, 3.0], I 29.8 [7.9, 3.1, 7.5, 8.7, 2.6], II 32.0 [8.7, 3.3, 8.3, 9.1, 2.6], III 26.8 [7.7, 2.8, 6.7, 7.4, 2.2], IV 29.1 [8.1, 2.7, 7.2, 8.7, 2.4].

Spination. Palp 131, 001, 1111; Legs: Femur I–III 323, IV 322; Patella I–IV 001/101; Tibia I–IV 2124/2224; Metatarsus I–III 2024, IV 3036.

Palp. As in diagnosis with weakly developed vRTA, ST huge and cymbium approximately twice as long as tibia (Figs 35a, b); ET wide and only distally narrowed, ET directed retrolaterad first and ending proximad at its distal end (Fig. 35c).

Female (n=2, single measurement: paratype):

Measurements. Medium sized. Total length 17.1; prosoma length 6.7, prosoma width 5.6,

anterior width of prosoma 3.6, opisthosoma length 10.4, opisthosoma width 7.5. Eye diameters:

AME 0.45, ALE 0.33, PME 0.31, PLE 0.33; eye interdistances: AME-AME 0.25, AME-ALE 0.13, PME-PME 0.45, PME-PLE 0.51, AME-PME 0.33, ALE-PLE 0.23, clypeus height at AME 0.35, clypeus height at ALE 0.45.

Chelicerae. Chelicerae dentition like as male.

Legs. Leg formula II I IV III. Measurements of palp and legs: Palp 7.5 [2.3, 1.1, 1.4, 2.7], I 23.7 [6.3, 2.9, 5.7, 6.7, 2.1], II 25.6 [7.0, 3.0, 6.2, 7.2, 2.2], III 20.0 [6.1, 2.2, 4.8, 5.3, 1.6], IV 22.3 [6.5, 2.5, 5.4, 6.3, 2.1].

Spination. Palp 131, 001, 1111, 1013; Legs: Femur I–III 323, IV 321; Patella I–IV 000/001; Tibia I–IV 2024; Metatarsus I–III 2024, IV 3036.

Epigyne/vulva. As in diagnosis with EF nearly as wide as long; MS thoroughly visible between lateral lobes (Fig. 36a); MS folded and in connection with extra membranous parts forming an inverse pocket visible in dorsal view between CDs (Fig. 36b); glandular pores restricted to a small depression near TL (Fig. 36c).

Colouration [in ethanol]. Yellowish cream prosoma with dark brown dorsal opisthosoma decorated with a chevron pattern, ventral opisthosoma pale in colour (Figs 55a–b).

Remarks. The copulatory structures of E. jocquei spec. nov. are strikingly different from other members of jaegeri group including the bifurcated dRTA and the distinctly visible MS between LL. However, it contains all the diagnostic characters of jaegeri species group. A female was found in ZMB, with label information ―Ostafrika‖ [=German for ―East Africa‖]. German East Africa used to be a German colony at the date of collecting (1904) which included what are now modern Tanzania, Rwanda, Mozambique and Burundi. Since the majority of the former German Results: chapter 3.2: Systematics and zoogeography with revision of Afro-Arabian species colony currently comprises the modern countries Tanzania and Mozambique, the specimen was probably collected from one of these countries.

Known geographical distribution and habitat. Known from the type locality and probably Tanzania or Mozambique in Southeast Africa.

tuckeri species group Diagnosis. Lacking intermarginal denticles in chelicerae (Figs 37e, 40e); female epigyne with AMLL fused together and encircling MS entirely (Figs 38a, 41a); male palps with an enlarged sub-tegulum (not as large as those of jaegeri-group) and very long embolus tip, embolus much longer relative to tegulum compared to other species groups; EM developed and forming a process covering ET partially (Figs 37a–c) to completely (Figs 40a–c).

Species composition. Two species: E. tuckeri comb. nov., E. educatus spec. nov.





Distribution. South-west Africa: Namibia and Angola (Fig. 71b).

–  –  –

Olios tuckeri Lawrence, 1927: 42, pl. 3, fig. 67 (description and illustration of male) [holotype ♂, examined]. New combination.

Olios furcatus Lawrence, 1927: 41, pl. 2, fig. 29 [syntype ♀ examined and designated as lectotype] [paralectotype ♂, undescribed genus] New synonymy (for justification see remarks).

Type material. Holotype of Olios tuckeri (designated by Lawrence 1927): male, NAMIBIA:

Kunene Region: Kunene River [label: Type, 1♂, Sparassidae, Eusparassus tuckeri lawr., South West Africa, Kunene R., c1712BC, R.F. Lawrence 1922, Shelf no. SAM/Aran 2639] (SAMC

B7124); Lectotype of Olios furcatus (designated here): female, NAMIBIA: Kunene Region:

Kunene River [label: South West Africa, Kunene R. c 1712BC, March 1923, R.F. Lawrence, Acc.no. B6625, Shelf no. SAM/Aran 2427] (SAMC B6625).

Results: chapter 3.2: Systematics and zoogeography with revision of Afro-Arabian species Other material examined. NAMIBIA: Kunene Region: 1♂, Epupa Falls, DK 334, 22 February 2005, D. Kunz leg. (SMF); 1♂, 1♀, 1sub♀, 1 juvenile, Etosha National Park, Sprokieswoud, 19˚ 05' S, 15˚ 37' E, 10 October 1986, under stones, E. Griffin leg. (NMNW 40564); 1♀, Etosha Pan, 18˚ 50' S, 16˚ 20' E, 4 March 1969, B. Lamoral & R. Day leg. (NMSA 12519); 2♂♂, Etosha National Park: Halali, 19˚ 01' S, 16˚ 29' E, 16 December 1993, B.M. Ullig leg. (ZMB); 2♀♀, SW of Windhoek, December 1929, R. Tucker leg.(SAMC 5162). Oshikoto Region: 2♀♀, Tsumeb, 1920, E. Koodig leg. (SAMC 4810). Otjozondjupa Region: 2♀♀, Road B8, S 19˚16.962', E 18˚26.935', 1233 m, 19 October 2009, M. Forman leg. (SD 802 & 803, SMF); 2♀♀, Grootfontein, 1919, R. M. Lightfoot leg. (SAMC 4625); ANGOLA: Namibe Province: 1♂, Parque Nacional de Iona (Iona National Park), 31 km S of Tombor, S 16˚20'36.1'', E 12˚26'21.1'', 241m, 11 January 2006, under stones, dense silk retreats, T. & C. Bird leg. (NMNW 45826); 1♀, 30–45 km NE of Namibe, 14˚ 55' 25.6'' S, 12˚ 22' 11.7'' E, 316 m, 12 January 2006, TB 06/36, T.

& C. Bird leg. (NMNW 45828); 1♀, Parque Nacional de Iona, 52 km NW of Espinheira, S 17˚04'26.9'', E 12˚03'40.9'', 564 m, 10 January 2006, dense silk retreats under stones, T. & C.

Bird leg. (NMNW 45829); 1♀, Iona district, ―Espaniera‖ (Espinheira), mountainous desert on grassy plain in between, 14 July 1996, R. Harris leg. (PPRI 96/595).

Diagnosis. Males easily distinguishable by long and slender ET directed proximad and by folded process of EM extending beyond ET retrolaterally (Figs 37a–c); females vulva with simple and straight CD and TL (Figs 38b–c, 39c) [compared to E. educatus spec. nov. with complicated vulvas] [see also diagnosis for tuckeri species group above].

Description. Male (ranges: n=6, single measurement: holotype):

Measurements (holotype first). Total length 8.7–10.1, prosoma length 4.2–5.1, prosoma width 3.5–4.7, anterior width of prosoma 1.8–2.6, opisthosoma length 4.5–5.0, opisthosoma width 3.0–

4.1. Eye diameters: AME 0.32, ALE 0.25, PME 0.23, PLE 0.27. Eye interdistances: AME-AME 0.20, AME-ALE 0.06, PME-PME 0.36, PME-PLE 0.30, AME-PME 0.24, ALE-PLE 0.20, clypeus height at AME 0.09, clypeus height at ALE 0.17. AME largest, ALE and PME subequal and smaller than PLE (Fig. 37d).

Chelicerae. Chelicerae with 2 anterior and 3 or 4 posterior teeth; cheliceral furrow without intermarginal denticles (Fig. 37e).

Results: chapter 3.2: Systematics and zoogeography with revision of Afro-Arabian species FIGURE 37. Eusparassus tuckeri (Lawrence, 1927) comb. nov., holotype male from Namibia: Kunene Riverside (SAMC). (a) left palp, ventral; (b) left palp, retrolateral; (c) embolus tip and conductor, ventral;

(d) eye arrangement, dorsal; (e) right chelicera, ventral.

Results: chapter 3.2: Systematics and zoogeography with revision of Afro-Arabian species FIGURE 38. Eusparassus tuckeri (Lawrence, 1927) comb. nov., female from Namibia: Kunene Riverside [lectotype female of ―Olios furcatus‖ (SAMC)]. (a) epigyne, ventral; (b) vulva, dorsal; (c) left vulva, anterio-dorso-lateral.

Results: chapter 3.2: Systematics and zoogeography with revision of Afro-Arabian species FIGURE 39. Eusparassus tuckeri (Lawrence, 1927) comb. nov., females from Angola (NMNW). (a, b) epigyne, ventral; (c) left vulva, anterio-dorso-lateral.

Legs. Leg formula: II IV=I III. Measurements of palp and legs: Palp 5.5 [1.7, 0.8, 0.9, 2.1], I 19.4 [5.5, 1.9, 5.1, 5.3, 1.6], II 23.2 [6.3, 2.2, 6.1, 6.7, 1.9], III 17.1 [5.0, 1.7, 4.1, 4.5, 1.8], IV 19.4 [5.5, 1.7, 4.9, 5.7, 1.6].

Spination. Palp 131, 001; Legs: Femur I–III 323, IV 322; Patella I–IV 101; Tibia I–IV 2124/2224; Metatarsus I–III 2024, IV 3034.

Palp. As in diagnosis with cymbium ~2.5 times longer than tibia; dRTA shortened and vRTA hump-like (Figs 37a, b).

Female (ranges: n=14, single measurement: lectotype):

Results: chapter 3.2: Systematics and zoogeography with revision of Afro-Arabian species Measurements (lectotype first). Total length: 12.2–14.3 prosoma length 4.6–6.1, prosoma width 3.9–5.0, anterior width of prosoma 3.0–3.3, opisthosoma length 7.6–8.2, opisthosoma width 4.0–

5.2. Eye diameters: AME 0.36, ALE 0.26, PME 0.25, PLE 0.31; eye interdistances: AME-AME 0.24, AME-ALE 0.05, PME-PME 0.42, PME-PLE 0.43, AME-PME 0.21, ALE-PLE 0.17, clypeus AME 0.15, clypeus ALE 0.20.

Chelicerae. Chelicerae with 2 anterior and 3 posterior teeth, cheliceral furrow with intermarginal denticles; one bristle at distal end of cheliceral basal segment.

Legs. Leg formula: II IV=I III. Measurements of palp and legs: Palp 5.4 [1.4, 0.7, 1.0, 2.3], I 16.5 [4.7, 2.1, 4.1, 4.2, 1.4], II 19.1 [5.4, 2.3, 4.8, 5.1, 1.5], III 14.0 [4.2, 1.7, 3.3, 3.4, 1.4], IV 16.9 [4.7, 2.2, 4.0, 4.6, 1.4].

Spination. Palp 131, 001, 1111, 1013; Legs: Femur I–III 323, IV 321/322; Patella I–IV 000/001;

Tibia I–IV 2024; Metatarsus I–III 2024, IV 3034.

Epigyne/vulva. As in diagnosis with EF slightly longer than wide (Fig. 38a), but in some specimens clearly longer than wide (Figs 39a, b); AMLL arch-shaped and LL enlarged laterally (Figs 38a, 39a, b); vulva with part of glandular pores shifted close to TL (Figs 38b, c; 39c).

Colouration [in ethanol]. Yellowish brown with uniform body colour in prosoma and legs;

dorsal opisthosoma with a darker band and surrounding dark patches (Fig. 56b).

Remarks. The type specimens were collected during the museum (SAMC) expedition to Kunene River in 1923. Lawrence (1927) described male and female of a single species under two different specific names. He classified them in the genus Olios. At that time, usage of the generic names Eusparassus and Olios was a subject of disputes, as the genera were not explicitly diagnosed. Simon (1932) was the first reviewer who distinguished between the genera Olios and Eusparassus (for more details see Moradmand and Jäger 2012b). However, this problem existed until recently as these two genera are very similar in terms of somatic characters and many transfers have been proposed (Moradmand & Jäger 2012a). Lawrence (1927) found two different male morphs and one female morph from the same locality, Kunene River. He described a male under the name tuckeri using a single male specimen. For some unexplained reason he described the female of tuckeri under the name furcatus along with a different male. This strange male (Lawrence 1927: fig. 68) belongs to a different, undescribed genus of Eusparassinae. According to the original description by Lawrence (1927), specific name furcatus is clearly assigned to the female. There are two main reasons for this judgment: first, Lawrence (1927) didnot list the accession number of the male (SAMC B6751) in the description, second he gave differential Results: chapter 3.2: Systematics and zoogeography with revision of Afro-Arabian species diagnosis just for the female. Thus, O. furcatus sensu Lawrence, 1927 is explicitly the female.

Accordingly, the female and male of O. furcatus are designated as lectotype and paralectotype, respectively. The female (lectotype of Olios furcatus) was misidentified by Lawrence and is clearly the conspecific female of E. tuckeri comb. nov. Finding of several sympatric males and females confirms this decision. Consequently, O. furcatus is proposed to be the junior synonym of E. tuckeri comb. nov. The paralectotype male of Olios furcatus resembles the genus Eusparassus in many somatic and genital characters but belongs to an undescribed genus. The vial of these specimens contained several male and also a female with a pre-epigyne, this female was probably overlooked by Lawrence.

Known geographical distribution and habitat. Northern Namibia and southern Angola (new country record) (Fig. 71b).

–  –  –

Type material. Holotype: male, NAMIBIA: Kunene Region: NE of Juriesdraai, Palmwag Lodge, under roof, S 19˚53.246', E 13˚56.203', 3 March 2005, DK 378, D. Kunz leg. (SMF).



Pages:     | 1 |   ...   | 18 | 19 || 21 | 22 |   ...   | 34 |


Similar works:

«A guide to immunisations up to one year of age Features the new MenB vaccination and the immunisation schedule from July 2016 the safest way to protect your child The complete immunisation programme from July 2016 is shown in detail in the back pages of this leaflet. Copies of these booklets are available from your clinic or doctor’s surgery. 2 A guide to immunisations up to one year of age Contents 4 Summary of immunisations up to one year of age 6 Common questions about immunisation 17 The...»

«Fun with Sounds Ideas for Small Group Work based on ‘Letters and Sounds Phase One’, with an emphasis on:  Tuning into sounds  Listening to and remembering sounds  Talking about sounds EARLY YEARS INCLUSION TEAM and PRE-SCHOOL FACILITATOR LETTERS AND SOUNDS Fun with Sounds Hopefully most people are now aware of the importance of developing good listening skills in children it is a strong element of ELKLAN, Letters and Sounds and EYFS training. To help you develop good listening...»

«BAKHTIN 81 Contre Bakhtin? The Problem of Poetry in Proust’s À la recherche du temps perdu JEFFERSON J. A. GATRALL, COLUMBIA UNIVERSITY Of the countless commentaries devoted to À la recherche du temps perdu, Proust’s own “Contre Sainte-Beuve” holds the unique distinction of having been written before the novel itself. At once a programmatic piece for a novel to come and a critique of biographical criticism, “Contre Sainte-Beuve” is an unfinished and heterogeneous text written, in...»

«MBIA Annual Report 2003 Capital Strength Triple-A Performance Pat Abt, Connie Accordino, Neil Ackerman, Joseph Adelizzi, Seth Albert, Angelique Allen, James Allen, Keith Allman, Mike Alter, Eric Anderson, Peter Andreou, Hector Angulo, John Antonazzo, Gina Antonelli, Brian Arcese, Ana Areal, John Ariola, Craig Armstrong, Errol Arne, Suresh Arumilli, Greer Avery, Dan Avitabile, Harold Bailey, Judy Baiocco, Michael Ballinger, Frances Banuelos, Ann Barbara, Mary Barbara, Lorraine Barg, Steve...»

«Distribution Agreement In presenting this thesis or dissertation as a partial fulfillment of requirements for an advanced degree from Emory University, I hereby grant to Emory University and its agents the non-exclusive license to archive, make accessible, and display my thesis or dissertation in whole or in part in all forms of media, now or hereafter known, including display on the world wide web. I understand that I may select some access restrictions as part of the online submission of this...»

«Capability Approach for Well-being Evaluation: A Case Study in Magelang Regency, Central Java, Indonesia R. Widodo D. Pramono (Department of Achitecture and Planning, Gadjah Mada University, Yogyakarta, Indonesia) Prof. Johan Woltjer (Faculty of Spatial Science, Groningen University, the Netherland) This paper is about research uses Amartya Sen’s Capability Approach (2000) to inductively devise a set of criteria for evaluating the well-being and quality of life, examining how the spatial...»

«AMERICAN ACADEMY OF ARTS AND SCIENCES GUIDE TO SERIES I-C-1: GENERAL RECORDS. COMMUNICATIONS TO THE ACADEMY (BOUND), 1780-1810. VOLUME 1, 1780-1790 VOLUME 2, 1785-1810 Archives American Academy of Arts and Sciences 136 Irving Street, Cambridge, MA 02138 © 2010 American Academy of Arts and Sciences Updated: 18 August 2010 AMERICAN ACADEMY OF ARTS AND SCIENCES ARCHIVES Series I-C-1: General Records. Communications to the Academy – Bound, 1780-1810 ADMINISTRATIVE INFORMATION Historical Note In...»

«Omaha, Nebraska Contractor/Trades Examination Information Bulletin PUBLISHED April 1, 2016 This edition supersedes all bulletin editions published prior to the above date. This bulletin answers most questions raised by examination candidates. Please read it carefully. You will find it a useful reference throughout your registration and examination process International Code Council Certification and Testing Department Birmingham District Office 900 Montclair Road Birmingham, AL 35213...»

«POWER OF ATTORNEY REVOCATION FORMS AND INSTRUCTIONS © Superior Court of Arizona in Maricopa County ALL RIGHTS RESERVED GNPOA_REV1 – 5336 101315 SELF-SERVICE CENTER REVOKE POWER OF ATTORNEY This packet contains court forms and instructions to file revoke power of attorney. Items in BOLD are forms that you will need to file with the Court. Non-bold items are instructions or procedures. Do not copy or file those pages! Order File Number Title # pages 1 GNPOA_REV1t Table of Contents (this page)...»

«A Middle East free of Missiles and Weapons of Mass Destruction: An Iranian View NASSER HADIAN Background paper Second EU Non-Proliferation Consortium Seminar to Promote Confidence Building and in Support of a Process Aimed at Establishing a Zone Free of WMD and Means of Delivery in the Middle East Brussels, 5-6 November 2012 The EU Non-Proliferation Consortium is a network of independent non-proliferation think tanks to encourage discussion of measures to combat the proliferation of weapons of...»

«Higher education and the Millennium Development Goals: where are we coming from, and where are we going? Liam Roberts and Patrice Ajai-Ajagbe October 2013 Published by The Association of Commonwealth Universities Woburn House 20-24 Tavistock Square London WC1H 9HF United Kingdom Tel: +44 (0)20 7380 6700 Fax: +44 (0)20 7387 2655 Email: beyond2015@acu.ac.uk www.acu.ac.uk/beyond-2015 © The Association of Commonwealth Universities 2013 While every care has been taken in the preparation of this...»

«Analiza Rynku Nieruchomości Miasta Turek 1. Turek w skrócie Turek miasto w województwie wielkopolskim, siedziba powiatu tureckiego i gminy Turek, w Kaliskiem, na ziemi sieradzkiej, na Wysoczyźnie Tureckiej, nad Kiełbaską. Według danych Urzędu Miasta stan na dzień 09.05.2013 wynosiła 28.655 mieszkańców. Miasto pełni funkcję ośrodka administracyjnego, przemysłowego, handlowo-usługowego i kulturalnego. W Turku próżno szukać idealnych zielonych łąk jak z reklamy, czy...»





 
<<  HOME   |    CONTACTS
2016 www.dissertation.xlibx.info - Dissertations, online materials

Materials of this site are available for review, all rights belong to their respective owners.
If you do not agree with the fact that your material is placed on this site, please, email us, we will within 1-2 business days delete him.